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Pre-publication versions of this article are available by contacting

Original Submission
14 Dec 2009 Submitted Original manuscript
10 Jan 2010 Reviewed Reviewer Report - Johannes Rieber
11 Jan 2010 Reviewed Reviewer Report - Felix Berger
11 Feb 2010 Reviewed Reviewer Report - Deniz Kececioglu
25 Feb 2010 Author responded Author comments - Julia Nuebel
Resubmission - Version 2
25 Feb 2010 Submitted Manuscript version 2
25 Mar 2010 Editorially accepted
25 Mar 2010 Article published 10.1186/1749-8090-5-15

How does Open Peer Review work?

Open peer review is a system where authors know who the reviewers are, and the reviewers know who the authors are. If the manuscript is accepted, the named reviewer reports are published alongside the article. Pre-publication versions of the article are available by contacting

You can find further information about the peer review system here.